Integrates information from physics (e.g., energetics), chemistry (the behavior of basic elements), and biology (evolutionary traits of species, multiple photosynthetic pathways, etc.) to understand the structure and functioning of ecosystems. Provides the background and necessary information to understand controls on photosynthesis, decomposition, and nutrient cycling across diverse terrestrial and aquatic landscapes. Prereqs.,EBIO 1210, 1220, 1230, 1240, and 2040, or EPOB 2050 and 2060, or EBIO 3020.